If insects rely on a negative temperature particle spinning around positive energy particles producing an anti-gravity effect for flight, then either the insects carries positive temperature particles or negative temperature particles. In the former case, the insect will seek low temperature ambiance places to fly in, and the latter high temperature ambiance place for flight. So, we have a hot wing and a cold wing. It is likely that a hot wing is hard in order to remain hot longer, and a cold wing is light and scaly to remain cool.
And so, hard winged insects (carrying positive temperature particle) seek low temperature environment and light scaly winged insects (carrying negative temperature particles) seek high temperature environment.
An electric field (electric static) applied to the elytra of a pood beetle, Dytiscidae, aligns/orientates the molecular structures holding the coupled pair of temperature particles and so directs the gravitational field in a specific direction.
The anti-gravitational effects of an elytra will disappear as ambient temperature raises.
Then, there are scaly butterfly wings.
Both types of wings are made of chitin; it is the micro structures on the wings that determine which type of temperature particles are in hold for controlled flight.
Speculatively yours.